Albanian non-marine molluscs

This is a complete list of non-marine molluscs from Albania, based on the 1996 list published by Dhimitër Dhora & Francisco Welter-Schultes.

Excerpt from

DHORA, DH. & WELTER-SCHULTES, F. W. (1996): List of species and atlas of the non-marine molluscs of Albania. -- Schriften zur Malakozoologie 9: 90-197. Cismar/Ostholstein.
 

Abstract of the "List of species and atlas":

The list of species of recent non-marine molluscs of Albania contains 292 species. 225 of them are land snails, 53 are freshwater Gastropoda and 14 are freshwater Bivalvia. Of these 292 species, 55 have only been mentioned in the literature without indication of certain localities where the species may live in Albania. For at least 33 of these species which have been mentioned in the literature, the occurence in Albania is doubtful. In the atlas, most of the literature references are included, new findings are added. Some of the most frequent and most important Albanian species are shown on plates.
Many of the species which are known from the non-Albanian parts of Lake Ohrid and Lake Shkodra may also occur in the Albanian parts of the lakes. A list of these species is given at the end of the atlas.

Lista e specieve të molusqeve të sotëm të tokës dhe të ujërave të ëmbla të Shqipërisë përmban 292 specie. 225 prej tyre janë gastropodë tokësorë, 53 janë gastropodë të ujërave të ëmbla dhe 14 janë bivalvë të ujërave të ëmbla. Prej këtyre 292 specieve, 55 vetëm sa përmenden në literaturë pa të dhëna për lokalitetet e caktuara se ku mund të jetojnë këto specie në Shqipëri. Për së paku 33 prej këtyre specieve, që përmenden në literaturë, dyshohet për ndodhjen në Shqipëri. Në atlas janë përfshirë shumica e referencave të literaturës, si dhe janë shtuar gjetjet e reja. Disa prej specieve më të shpeshta dhe më të rëndësishme të Shqipërisë paraqiten në fletët ilustruese.
Disa prej specieve që njihen prej pjesëve jo shqiptare të Liqenit të Ohrit dhe Liqenit të Shkodrës mundet që të ndodhen edhe në pjesët shqiptare të liqeneve. Në fund të atlasit jepet një listë e këtyre specieve.
 
 

Geographic limit of this work

Most species described from the non-Albanian parts of Lake Ohrid may also occur in the Albanian parts of the lake. Of course there is not much sense in publishing an atlas including the Albanian parts of Lake Shkodra, Lake Ohrid and the other lakes, leaving out the non-Albanian parts. We do not know much about the molluscs of other regions, but the molluscs in the Balkans are generally not interested in where the actual national borders of human administrative areas are. However, a geographic limit of this work had to be chosen somewhere, so we finally decided to follow the national borders. For Lake Ohrid and Lake Shkodra, lists of species are given after the atlas.
 

List of species

In the list of species, the systematic order of families presented in FALKNER 1990 is followed. Systematics in SE-Europe is much more difficult than it is in Central and Northern Europe. There are more species per area, more trends to micro-endemisms, more species are divided into subspecies, and everything is less well known. Changes in taxonomy are much more frequent than in Central Europe. Some groups have been revised recently, some groups are about to be revised in the present years, and other groups are still "waiting" for their "specialist" falling in love with them. The list of species reflects our actual knowledge on non-marine molluscs in Southern Europe in general.
The systematic position of many described "species" is not known at all. An example for this situation is Helicigona "albanica" S. H. F. JAECKEL 1954. For systematical statements in Helicigona anatomical dissection is necessary, and there is no conserved material. The access to the type locality is very difficult and scientists should not go there alone. Finally, if it is a "1954 new" species in the genus Helicigona, its name must be changed (because of albanica KOBELT 1902).
 

Syn./ssp.: Synonymy, subspecies

In many cases it is very difficult to decide whether we deal with a subspecies or a synonym. In most groups of molluscs in SE-Europe, we still do not know enough for solid statements on subspecies level. In the low number of points in the atlas, compared with the much higher number of points in equal works for Central or Northern European regions, this situation of very poor knowledge cannot be reflected more clearly.
The percentage of subspecies which have been reported from Albania and which certainly do not live in the country is high, probably much higher than it is in the case of species. These are the reasons why we decided to put synonyms and subspecies in one box.
 

List of non-marine mollusc species of Albania

Syn./ssp. = synonymy and/or subspecies
 

Prosobranchia: Archaeogastropoda
Neritoidea
Neritidae
Former generic names: Neritina, Theodoxia.
Family systematics: according to FALKNER 1990.

Theodoxus fluviatilis (LINNÉ 1758)
Syn./ssp.: dalmatina, danubialis C. PFEIFFER 1828, scutariensis
Note: Th. danubialis does not live in today's Albania.

Theodoxus varius (MENKE 1828)
 

Caenogastropoda
Cyclophoroidea
Cochlostomatidae
Former generic names: Auritus, Pomatias, Pomatius.
Family systematics: according to SCHÜTT 1977.
 
 

Ampullarioidea
Viviparidae
Former generic names: Contectiana, Cyclostoma, Paludina, Vivipara.
Family systematics: according to FALKNER 1990.

Viviparus contectus (MILLET 1813)

Viviparus janinensis (MOUSSON 1859)
Note: Occurence in Albania needs confirmation.

Viviparus mamillatus (KÜSTER 1852)

Viviparus viviparus (LINNÉ 1758)
 

Valvatoidea
Valvatidae

Valvata cristata O. F. MÜLLER 1774
Note: Lives in Shkodra Lake.

Valvata piscinalis (O. F. MÜLLER 1774)

Valvata rhabdota STURANY 1894
Note: Occurence in Albania needs confirmation.
 

Littorinoidea
Pomatiasidae
Former generic names: Cyclostoma, Ericia.

Pomatias elegans (O. F. MÜLLER 1774)
 

Hydrobioidea (= Truncatelloidea)
Hydrobiidae
Former generic names: Orientalia, Prespolitoralia.
Family systematics: according to RADOMAN 1983.
Obviously, the international scientific community did not follow RADOMAN (1983) in all questions of generic names of Hydrobioidea. In our list, the generic names generally follow RADOMAN 1983, but the division in separate families (like Orientalinidae etc.) is not adopted.
 

Sadleriana virescens (KÜSTER 1852)
Note: Systematic position not clear, possibly = Orientalina curta.

Horatia klecakiana BOURGUIGNAT 1887
Syn./ssp.: albanica
Note: Lives probably not in today's Albania.

Adriohydrobia consociella (FRAUENFELD 1863)
Note: Occurence in Albania needs confirmation.

Emmericia patula (BRUMATI 1838)
Note: Occurence in Albania needs confirmation.

Hydrobia ventrosa (MONTAGU 1803)

Orientalina curta (KÜSTER 1852)
Syn./ssp.: albanica RADOMAN 1973[a], kicavica

Anagastina scutarica (RADOMAN 1973[b])
Note: Lives in Shkodra Lake.

Chilopyrgula sturanyi (BRUSINA 1896)

Ohridopyrgula macedonica (BRUSINA 1896)

Ohridohoratia pygmaea (WESTERLUND 1902)

Zaumia kusceri (HADZISCE 1956)

Polinskiola polinskii (RADOMAN 1960)

Ohrigocea stankovici (HADZISCE 1956)

Albaniana albanica RADOMAN 1973[a]

Prespolitorea malaprespensis (RADOMAN 1973[a])
Syn./ssp.: malaprespaensis

Malaprespia albanica RADOMAN 1973[a]

Parabythinella malaprespensis RADOMAN 1973[a]
Syn./ssp.: malaprespaensis
 Pyrgula annulata (LINNÉ 1758)
 

Bithyniidae
Former generic names: Bythinia.
Family systematics: according to SCHÜTT 1987, SCHÜTT 1988.
 

Bithynia tentaculata (LINNÉ 1758)
Syn./ssp.: mostarensis, majewskyi
 

Aciculidae
Former generic names: Acicula, Acme.
Family systematics: according to BOETERS & al. 1989. The list has been checked by E. GITTENBERGER (Leiden), 1995.

Platyla banatica (ROSSMÄSSLER 1842)

Platyla pinteri SUBAI 1976

Platyla wilhelmi (A. J. WAGNER 1910)

Renea kobelti (A. J. WAGNER 1910)
Syn./ssp.: albanica BOETERS & al. 1989
 

Cerithioidea
Thiaridae
Former generic names: Amphimelania, Melania.

Holandriana holandrii (C. PFEIFFER 1828)
Syn./ssp.: laevigata
 

Pulmonata: Basommatophora
Ellobioidea
Carychiidae (= Ellobiidae)

Ovatella myosotis (DRAPARNAUD 1805)
Note: Occurence in Albania needs confirmation.
 

Acroloxoidea
Acroloxidae

Acroloxus lacustris (LINNÉ 1758)
 

Planorboiodea
Physidae

Aplexa hypnorum (LINNÉ 1758)
 
Physa fontinalis (LINNÉ 1758)
Syn./ssp.: dalmatina

Physella acuta (DRAPARNAUD 1805)
 

Planorbidae
Family systematics: according to HUBENDICK 1970, MEIER-BROOK 1983.
 

Ancylus fluviatilis O. F. MÜLLER 1774
Syn./ssp.: capuloides, recurvus

Anisus vortex (LINNÉ 1758)

Anisus vorticulus (TROSCHEL 1834)

Bathyomphalus contortus (LINNÉ 1758)
 
Ferrissia wautieri (MIROLLI 1960)

Gyraulus albus (O. F. MÜLLER 1774)

Gyraulus crenophilus HUBENDICK & RADOMAN 1959

Gyraulus crista (LINNÉ 1758)

Gyraulus fontinalis HUBENDICK & RADOMAN 1959

Gyraulus laevis (ALDER 1838)

"Gyraulus plavensis A. J. W[AGNER]."
Note: Systematic position not clear, probably the taxon has never been described.
 
Planorbarius corneus (LINNÉ 1758)

Planorbis carinatus O. F. MÜLLER 1774

Planorbis planorbis (LINNÉ 1758)
 

Lymnaeoidea
Lymnaeidae
Former generic names: Gulnaria, Limnaea, Limnophysa.
Family systematics: according to FALKNER 1990.

Galba truncatula (O. F. MÜLLER 1774)

Lymnaea stagnalis (LINNÉ 1758)

Radix ampla (HARTMANN 1821)
 
Radix auricularia (LINNÉ 1758)

Radix ovata (DRAPARNAUD 1805)

Radix peregra (O. F. MÜLLER 1774)
 
Radix relicta POLINSKI 1929

Stagnicola corvus (GMELIN 1791)

Stagnicola palustris (O. F. MÜLLER 1774)
 

Stylommatophora
Cochlicopoidea
Cochlicopidae
Family systematics: The list has been checked by E, GITTENBERGER (Leiden), 1995.

Cochlicopa lubrica (O. F. MÜLLER 1774)

Cochlicopa lubricella (PORRO 1838)

Hypnophila polita (PORRO 1838)
Note: Lives very probably in South Albania.

Hypnophila pupaeformis (CANTRAINE 1835)
 

Pupilloidea
Pyramidulidae
Family systematics: The list has been checked by E, GITTENBERGER (Leiden), 1995.

Pyramidula cephalonica (WESTERLUND 1898)

Pyramidula pusilla (VALLOT 1801)
 
 

Chondrinidae
Former generic names: Abida, Modicella, Pupa, Torquilla.
Family systematics: according to NORDSIECK 1970b, GITTENBERGER 1973, GITTENBERGER 1984. The list has been checked by E, GITTENBERGER (Leiden), 1995.
 

Chondrina arcadica (REINHARDT 1881)
Syn./ssp.: clienta WESTERLUND 1883
Plate 7 Fig. 56-57

Chondrina spelta (BECK 1837)
Syn./ssp.: Pupa muehlfeld(ti)i KÜSTER 1843, fuchsi
Plate 7 Fig. 60-61

Granaria frumentum (DRAPARNAUD 1801)
Syn./ssp.: illyrica ROSSMÄSSLER 1837
Plate 7 Fig. 62-63

Granopupa granum (DRAPARNAUD 1801)
Plate 7 Fig. 58-59

Rupestrella philippii (CANTRAINE 1840)
Plate 7 Fig. 52-53

Rupestrella rhodia (ROTH 1839)
Plate 7 Fig. 54-55
 

Argnidae
Former generic names: Argna, Agardhia,, Coryna.
Family systematics: The list has been checked by B. HAUSDORF (Hamburg), 1996.

Agardhiella truncatella (L. PFEIFFER 1841)
Syn./ssp.: skipetarica [sic!] A. J. WAGNER 1914
 

Orculidae
Family systematics: The list has been checked by B. HAUSDORF (Hamburg), 1996.

Orcula schmidtii (KÜSTER 1843)
Syn./ssp.: transversalis
Plate 7 Fig. 65

Orcula wagneri STURANY 1914
Plate 7 Fig. 64

Pagodulina gracilior PILSBRY 1926
Syn./ssp.: gracilis WESTERLUND 1887 non ROSSMÄSSLER 1842
Plate 7 Fig. 67-68

Pagodulina subdola (GREDLER 1856)
Syn./ssp.: kaeufeli KLEMM

Sphyradium doliolum (BRUGUIERE 1792)
Plate 7 Fig. 66
 

Lauriidae
Former generic names: Charadrobia.
Family systematics: The systematic position of this group has been checked by B. HAUSDORF (Hamburg), 1996.

Lauria cylindracea (DA COSTA 1778)
Syn./ssp.: umbilicus ROTH 1839
 

Pupillidae
Former generic names: Aspasita.
Family systematics: The list has been checked by E, GITTENBERGER (Leiden), 1995.

Pupilla alpicola (CHARPENTIER 1837)

Pupilla triplicata (STUDER 1820)

Spelaeodiscus albanicus (A. J. WAGNER 1914)
 

Valloniidae
Former generic names: Planogyra.
Family systematics: The list has been checked by E, GITTENBERGER (Leiden), 1995.

Acanthinula aculeata (O. F. MÜLLER 1774)
Note: Occurence in Albania needs confirmation.

Gittenbergia sororcula (BENOIT 1857)
Note: Occurence in Albania needs confirmation.

Vallonia costata (O. F. MÜLLER 1774)

Vallonia enniensis (GREDLER 1856)
Syn./ssp.: costellata SANDBERGER

Vallonia excentrica STERKI 1982

Vallonia pulchella (O. F. MÜLLER 1774)
 

Enidae = Buliminidae
Former generic names: Buliminus, Bulimus, Napaeus, Zebrinus.
Family systematics: The list has been checked by E, GITTENBERGER (Leiden), 1995.
Genus Ena, Zebrina, Napaeopsis: BANK & MENKHORST 1992
 
Chondrula consentanea KÜSTER 1870
Note: Occurrence in Albania needs confirmation.

Chondrula lugorensis A. J. WAGNER 1914

Chondrula macedonica A. J. WAGNER 1914

Chondrula microtraga (ROSSMÄSSLER 1839)

Chondrula tridens (O. F. MÜLLER 1774)
Syn./ssp.: eximia, quinquedentata

Ena concolor (WESTERLUND 1887)
Syn./ssp.: jugoslaviensis

Ena montana (DRAPARNAUD 1801)

Ena subtilis (ROSSMÄSSLER 1835)
Syn./ssp.: reitteri

Imparietula seductilis (ROSSMÄSSLER 1837)

Jaminia quadridens (O. F. MÜLLER 1774)
Syn./ssp.: nicollii, Chondrula priamus (KOBELT 1893) sensu GAMBETTA 1930

Mastus pupa (LINNÉ 1758)
Syn./ssp.: pupa BRUGUIERE 1792

Merdigera obscura (O. F. MÜLLER 1774)

Napaeopsis cefalonicus (MOUSSON 1859)
Syn./ssp.: merditanus STURANY 1907, zebianus, latifianus, winneguthi
Note: Polytypic species.

Zebrina detrita (O. F. MÜLLER 1774)
Syn./ssp.: inflata, radiata, tumida, tumidus
 

Clausilioidea
Clausiliidae
Former generic names: Alinda, Alopia, Clausiliastra, Laciniaria, Pirostoma, Uncinaria.
Family systematics: according to NORDSIECK 1979.
Genus Agathylla: NORDSIECK 1972, NORDSIECK 1996
Genus Albinaria: NORDSIECK 1974, 1977a
Genus Cochlodina: GITTENBERGER 1967
Genus Delima: NORDSIECK 1969
Genus Herilla: NORDSIECK 1971
Genus Medora: NORDSIECK 1970a
Genus Montenegrina: NORDSIECK 1972, 1974, 1977b
Genus Protoherilla: NORDSIECK 1972
Genus Triloba: NORDSIECK 1972

Herilla bosniensis (L. PFEIFFER 1868)
Syn./ssp.: ibarensis, semistriata, gastron, exornata, plivae, travnicana, brandisi

Herilla illyrica (MÖLLENDORFF 1899)
Syn./ssp.: miosis, diabasis, medoroides, vidascens, oribates

Herilla jabucica BÖTTGER [= O. BOETTGER] 1907 [in WOHLBEREDT 1907a]
Syn./ssp.: albanica S. H. F. JAECKEL 1956, iabucica, jabukica
Note: iabucica = printing error for jabucica in WOHLBEREDT 1907a: 550.

Herilla ziegleri (KÜSTER 1845)
Syn./ssp.: klemmi

Triloba sandrii (KÜSTER 1844)
 Triloba thaumasia (STURANY 1907)
Syn./ssp.: korabensis, dardanorum, faueri

Medora proxima (WALDERDORFF 1864)
Syn./ssp.: remota POLINSKI 1924
Note: In Albania only M. p. remota.
 

Isabellaria lophauchena (STURANY 1894)
Note: Does not live in today's Albania, only in Greece.

Albinaria contaminata (ROSSMÄSSLER 1835)
Syn./ssp.: artaensis, fuscostrigata, lactea, muraria, odysseus
Notes: artaensis refers to Arta in Ipiros, not in today's Albania. A. contaminata does not live in today's Albania (NORDSIECK 1974: 134).

Albinaria hians (O. BOETTGER 1878 [b])
Note: Does not live in today's Albania, only in Greece.

Albinaria schuchii (ROSSMÄSSLER 1836)

Albinaria scopulosa (CHARPENTIER 1852)
Syn./ssp.: winneguthi, epirotes

Strigilodelima conspersa (L. PFEIFFER 1848)
Syn./ssp.: hypermegala, invalida, platystoma KÜSTER 1853

Strigilodelima pentheri (A. J. WAGNER 1919)

Montenegrina apfelbecki (STURANY 1907)

Montenegrina cattaroensis (ROSSMÄSSLER 1835)
Syn./ssp.: gracilior DESHAYES
Note: Does not live in today's Albania.

Montenegrina dofleini (A. J. WAGNER 1928)
Syn./ssp.: kaiseri BRANDT 1961

Montenegrina ersekensis NORDSIECK 1996

Montenegrina fuchsi BRANDT 1961
Syn./ssp.: klemmi

Montenegrina helvola (KÜSTER 1860)
Syn./ssp.: pageti

Montenegrina irmengardis KLEMM 1962

Montenegrina janinensis (MOUSSON 1859)
Syn./ssp.: attemsi,coerulescens, fagorum, tomorosi

Montenegrina laxa (KÜSTER 1861)
Syn./ssp.: chiasma, iba, miraka, weigneri

Montenegrina perstriata (A. J. WAGNER 1919)
Syn./ssp.: scipetarica, skipetarica SOOS 1924
 

Montenegrina umbilicata (O. BOETTGER 1879)
Syn./ssp.: costata O. BOETTGER 1907
Note: Does not live in today's Albania (NORDSIECK 1972: 27)

Protoherilla baleiformis (O. BOETTGER 1909)
Syn./ssp.: janicki(i) A. J. WAGNER 1921

Protoherilla pseudofallax (H. NORDSIECK 1972)

Cochlodina laminata (MONTAGU 1803) (Cochlodina)
Syn./ssp.: albanica S. H. F. JAECKEL 1956, nana MÖLLENDORFF 1873, oreinos, tertia O. BOETTGER 1907
Note: It is not sure if all reports in literature refer to this species.

Delima bilabiata (SCHUBERTH & WAGNER 1829) (Semirugata)
Syn./ssp.: alschingeri

Delima binotata (ROSSMÄSSLER 1836)
Syn./ssp.: binodata, gastrolepta
Note: Does not live in today's Albania (NORDSIECK 1969: 271).

Delima montenegrina (L. PFEIFFER 1848) (Delima)
Syn./ssp.: digamma O. BOETTGER 1878 [a], inermis, muralis, nodulosa, pseudobinodata, semilabiata, tabida

Siciliaria pantocratoris (O. BOETTGER 1889)

Siciliaria stigmatica (ROSSMÄSSLER 1836) (Stigmatica)
Syn./ssp.:margaritifera, maritima, splendens, sturmi, sturmii L. PFEIFFER 1848

Papillifera papillaris (O. F. MÜLLER 1774)
Syn.: bidens LINNÉ

Clausilia dubia DRAPARNAUD 1805 (Andraea)
Syn./ssp.: transsilvanica A. SCHMIDT 1857
Note: Probably in Albania, but needs confirmation.

Balea biplicata (MONTAGU 1803) (Alinda)
Syn./ssp.: distincta, eupleuris, labiata
Note: B. p. distincta does not live in Albania.

Balea serbica MÖLLENDORFF 1873
Syn./ssp.: fallax ROSSMÄSSLER 1836
Note: Probably in Albania, but still needs confirmation.

Vestia roschitzi (BRANCSIK 1889) (Vestiella)
Syn./ssp.: pygmaea, apragmosyne

Bulgarica thessalonica (ROSSMÄSSLER 1839)
Note: Does not live in Albania.

Bulgarica vetusta (ROSSMÄSSLER 1836) (Strigilecula)
Syn./ssp.: striolata
 

Succineoidea
Succineidae

Oxyloma elegans (RISSO 1826)

Succinea oblonga (DRAPARNAUD 1801)

Succinea pfeifferi ROSSMÄSSLER 1835
 

Achatinoidea
Ferussaciidae

Cecilioides acicula (O. F. MÜLLER 1774)

Cecilioides tumulorum (BOURGUIGNAT 1856)

Cecilioides spec. 3
Note: Cecilioides spec. 3 is an intermediate form between C. acicula and C. tumulorum, possibly a bastard form. The systematics in Cecilioides is still not clear.
 

Subulinidae

Rumina decollata (LINNÉ 1758)
Note: Occurence in Albania needs confirmation.
 

Testacelloidea
Oleacinidae
Former generic names: Glandina, Oleacina.
Family systematics: according to SUBAI 1980. The list has been checked by P. SUBAI (Aachen), 1995.

Poiretia compressa (MOUSSON 1859)

Poiretia cornea (BRUMATI 1838)

Poiretia delesserti (BOURGUIGNAT 1852)
 

Punctoidea
Punctidae

Punctum pygmaeum (DRAPARNAUD 1801)
 

Gastrodontoidea
Gastrodontidae
Family systematics: The list has been checked by A. RIEDEL (Warszawa), 1995.

Zonitoides nitidus (O. F. MÜLLER 1774)
 

Euconuloidea
Euconulidae

Euconulus fulvus (O. F. MÜLLER 1774)
 

Vitrinoidea, [Zonitoidea]
Vitrinidae
Former generic names: Phenacolimax, Semilimacella.
Family systematics: The list has been checked by B. HAUSDORF (Hamburg), 1995.

Gallandia annularis (STUDER 1820)

Oligolimax reitteri (O. BOETTGER 1880)

Semilimax apatelus (SOOS 1924)
Note: Systematic position not clear.

Vitrina carniolica O. BOETTGER 1884 (Oligolimax ?)
Note: Systematic position not clear.

Vitrina pellucida (O. F. MÜLLER 1774)
Note: Occurence in Albania needs anatomical confirmation.
 

Zonitidae
Former generic names: Aegopina, Crystallus, Helix, Hyalina, Hyalinia, Hydatinus, Morlina, Polita, Tragomma, Zonites.
Family systematics: The list has been checked by A. RIEDEL (Warszawa), 1995.
Genus Allaegopis: RIEDEL 1982, RIEDEL 1993b
Genus Gyralina: RIEDEL & SUBAI 1993
Genus Paraegopis: RIEDEL 1978b
Genus Vitrea: PINTÉR 1972

Vitrea botterii (L. PFEIFFER 1853)

Vitrea contracta (WESTERLUND 1871)
Syn./ssp.: zakynthia

Vitrea illyrica (A. J. WAGNER 1907)

Vitrea selecta PINTÉR 1972
Note: Systematic position is not clear yet. Belongs possibly to V. botterii.

Vitrea sturanyi (A. J. WAGNER 1907)
Note: Occurence in Albania needs confirmation.

Vitrea subrimata (REINHARDT 1871)
Syn./ssp.: inflata A. J. WAGNER 1907, sphaerocona

Gyralina candida (A. J. WAGNER 1909 [in WOHLBEREDT 1909a])
Syn./ssp.: Zonitoides candidus

Gyralina circumlineata (L. PFEIFFER 1846)

Gyralina gjirokastrana RIEDEL & WELTER-SCHULTES 1996

Gyralina korabensis (RIEDEL 1970)
Note: Lives probably in Albania.

Gyralina tarabosensis (RIEDEL 1970)

Aegopinella minor (STABILE 1864) [s. l.]
Syn./ssp.: inermis A. J. WAGNER 1907 (not sensu FORCART 1959, = A. forcarti), nitens
Note: A. nitens certainly does not live in Albania.

Allaegopis amphikypellon RIEDEL 1982

Allaegopis skanderbegianus (POLINSKI 1924)
Syn./ssp.: polinskii URBANSKI, ljubetinensis

Allaegopis transiens (MOUSSON 1859)
Note: Occurence in Albania needs confirmation.

Paraegopis albanicus (ROSSMÄSSLER 1836)

Paraegopis bizonus A. J. WAGNER 1914

Paraegopis mauritii (WESTERLUND 1886)
Syn./ssp.: montenegrinus O. BOETTGER [in WOHLBEREDT] 1909
Note: Occurence in Albania needs confirmation.

Paraegopis skipetaricus A. J. WAGNER 1914 (?)
Note: Taxonomic revision is needed.

Aegopis acies (FÉRUSSAC 1832)
Note: Occurence in Albania needs confirmation. Taxonomy of Aegopis and distribution of its species is not cleared yet.

Aegopis septentrionalis laughofferi A. J. WAGNER 1914
Note: Taxonomy of Aegopis and distribution of its species is not cleared yet.

Aegopis verticillus (LAMARCK 1822)
Note: Occurence in Albania needs confirmation. Taxonomy of Aegopis and distribution of its species is not cleared yet.

Oxychilus camelinus (BOURGUIGNAT 1852)

Oxychilus cellarius (O. F. MÜLLER 1774)
Note: Lives probably not in today's Albania.

Oxychilus cyprius (L. PFEIFFER 1847)
Syn.: macedonicus RIEDEL 1959

Oxychilus depressus (STERKI 1880)

Oxychilus draparnaudi (BECK 1837)
Note: Occurence in Albania is anatomically confirmed.

Oxychilus glaber (ROSSMÄSSLER 1835)
Syn./ssp.: bojanae A. J. WAGNER 1907, glabra, nitidissima, nitidissimus

Oxychilus hydatinus (ROSSMÄSSLER 1838)
Syn./ssp.: eudedalaea BOURGUIGNAT

Oxychilus planorbis (MÖLLENDORFF 1899)/ Oxychilus juliae RIEDEL 1990 [complex]
 

Daudebardiidae
Family systematics: according to RIEDEL 1978a. The list has been checked by A. RIEDEL (Warszawa), 1995.

Carpathica stussineri (A. J. WAGNER 1895)

Daudebardia brevipes (DRAPARNAUD 1805)

Daudebardia rufa (DRAPARNAUD 1805)
Syn./ssp.: cycladum
 

Milacidae
Former generic names: Amalia, Limax, Milax.
Family systematics: according to WIKTOR 1987. The list has been checked by A. WIKTOR (Wroclaw), 1995.

Tandonia albanica (SOOS 1924)
Syn./ssp.: adelpha, clerxi

Tandonia sowerbyi (A. FÉRUSSAC 1823)
 

Limacoidea
Limacidae
Family systematics: The list has been checked by A. WIKTOR (Wroclaw), 1995.

Gigantomilax csikii SOOS 1924
Note: This species certainly does not belong to the genus Gigantomilax, but more probable to Deroceras (Agriolimacidae).

Lehmannia marginata (O. F. MÜLLER 1774)

Lehmannia szigethyae WIKTOR 1975

Limacus flavus (LINNÉ 1758)

Limax carbonarius O. BOETTGER 1885
Syn./ssp.: (albanicus JAECKEL 1954)
Note: This species is probably conspecific with Limax cephalonicus SIMROTH 1886.

Limax cinereoniger WOLF 1803
Syn./ssp.: vera

Limax maximus LINNÉ 1758
Syn./ssp.: cinereus, (albanicus JAECKEL 1954)

Limax wohlberedti SIMROTH 1900
 

Agriolimacidae
Former generic names: Agriolimax.
Family systematics: The list has been checked by A. WIKTOR (Wroclaw), 1995.
Note: Gigantomilax csikii SOOS 1924 (see Limacidae) could belong to the genus Deroceras (Agriolimacidae).

Deroceras agreste (LINNÉ 1758)

Deroceras reticulatum (O. F. MÜLLER 1774)

Deroceras sturanyi (SIMROTH 1894)
Note: Occurence in Albania needs confirmation.

Deroceras turcicum (SIMROTH 1894)
 

Arionoidea
Arionidae
Family systematics: The list has been checked by A. WIKTOR (Wroclaw), 1995.

Arion circumscriptus JOHNSTON 1828

Arion rufus (LINNÉ 1758)
Note: Lives probably not in today's Albania.

Arion subfuscus (DRAPARNAUD 1805)
 

Helicoidea
Hygromiidae
Former generic names: Anchistoma, Ashfordia, Bradybaena, Caracollina, Carthusiana, Eulota, Fruticicola, Gonostoma, Helicella, Helicopsis, Heliomanes, Helix, Hygromia, Metafruticicola, Polygyra, Theba, Trigonostoma, Trochula, Vortex, Xerophila, Zenobia.
Family systematics: according to NORDSIECK 1993. The list has been checked by B. HAUSDORF (Hamburg), 1995.
 

Candidula castriota SOOS 1924
Note: Systematic position not clear.

Cernuella virgata (DA COSTA 1778)
Syn./ssp.: variabilis DRAPARNAUD 1805, variegata

Cochlicella acuta (O. F. MÜLLER 1774)
Syn./ssp.: Bulimus acutus

Cochlicella barbara (LINNÉ 1758)

Helicodonta albanica A. J. WAGNER 1914

Helicodonta langhofferi A. J. WAGNER 1912
Syn./ssp.: laughofferi
Note: Lives probably not in Albania.

Helicodonta obvoluta (O. F. MÜLLER 1774)

Lindholmiola lens (FÉRUSSAC 1832)

Lindholmiola girva (FRIVALDSZKY 1835)
Syn./ssp.: canalifera, contorta ROSSMÄSSLER 1838, corcyrensis ROSSMÄSSLER 1838, octogyrata, pirinensis

Hiltrudia mathildae (WESTERLUND 1881)
Syn./ssp.: kusmici CLESSIN 1887

Monacha cartusiana (O. F. MÜLLER 1774)-complex
Note: Contains M. cartusiana and M. dissimulans, which can only be determined by anatomical dissection.

Monacha cartusiana (O. F. MÜLLER 1774)
Syn./ssp.: carthusiana

Monacha dissimulans PINTÉR 1968
Note: There are probably older names for M. dissimulans.

Monacha frequens (MOUSSON 1859)-complex
Note: It is not clear if the Albanian M. frequens are conspecific. Reports of M. parumcincta probably refer to M. frequens-complex.

Monacha microtricha S. H. F. JAECKEL 1954
Note: Systematic position is not clear.

Monacha parumcincta (MENKE 1828) [not ROSSMÄSSLER 1837]
Syn./ssp.: olivieri
Note: Occurence in Albania needs confirmation. Reports in literature refer probably to M. frequens-complex.

Monachoides incarnatus (O. F. MÜLLER 1774)

Petasina erjaveci (BRUSINA 1870)
Syn./ssp.: floerickei (KOBELT 1898)

Pseudotrichia rubiginosa (ROSSMÄSSLER 1838)
Syn./ssp.: epirotica

Semifruticicola serbica A. J. WAGNER 1914
Syn./ssp.: costulata BRANCSIK 1897
Note: Systematic position not clear.

Trichia hispida (LINNÉ 1758)
Note: Lives most probably not in today's Albania.

Trochoidea pyramidata (DRAPARNAUD 1805)

Trochoidea trochoides (POIRET 1789)
Syn./ssp.: conica DRAPARNAUD, crenulata GERMAIN

Xerolenta obvia (MENKE 1828)
Syn./ssp.: arenosa, candicans
Note: Reports for South and Central Albania refer most probably to Xeromunda vulgarissima.

Xeromunda vulgarissima (MOUSSON 1859)

Xeropicta derbentina (KRYNICKI 1836)
Syn./ssp.: gyroides, homoleuca

Xerotricha conspurcata (DRAPARNAUD 1801)
 

Helicidae
Ariantinae
Former generic names for Ariantinae: Arianta, Campylaea, Cattania, Chilostoma, Cingulifera, Corneola, Dinarica, Faustina, Liburnica, Helix, Iberus.
Subfamily systematics: The list for Ariantinae has been checked by P. SUBAI (Aachen), 1995.

Helicigona albanica S. H. F. JAECKEL 1954 [not KOBELT 1902]
Note: Systematic position not clear.

Helicigona albanograeca SUBAI 1995
Note: Lives probably in South Albania.

Helicigona apfelbecki bindzaensis (STURANY 1909)
Syn./ssp.: braueri A. J. WAGNER 1914, prokletijensis KNIPPER 1939

Helicigona byshekensis KNIPPER 1941

Helicigona denudata (ROSSMÄSSLER 1836)
Note: Does not live in today's Albania.

Helicigona dochii (STURANY 1907)
Syn./ssp.: munelana

Helicigona dunjana KNIPPER 1941

Helicigona edlaueri KNIPPER 1941

Helicigona fuchsi KNIPPER 1939

Helicigona harpya SUBAI 1995

Helicigona hoffmanni (ROSSMÄSSLER 1836)
Syn./ssp.: walteri

Helicigona inflata (KOBELT 1876)
Syn./ssp.: rumelica, trizona ROSSMÄSSLER 1835
Note: H. trizona has to be considered as a distinct species.

Helicigona insolita (ROSSMÄSSLER 1838)
Syn./ssp.: insolida
Note: Does not live in today's Albania.

Helicigona maranajensis (A. J. WAGNER 1914)

Helicigona petrovici (A. J. WAGNER 1914)

Helicigona pouzolzii (DESHAYES 1830) [not PAYRAUDEAU 1826]
Syn./ssp.: albanica KOBELT 1902, bosniensis, montenegrina ROSSMÄSSLER 1836, pouzolzi

Helicigona reischuetzi (SUBAI 1990)

Helicigona serbica (KOBELT 1872)

Helicigona setigera (ROSSMÄSSLER 1836)

Helicigona setosa (ROSSMÄSSLER 1836)
Note: Lives certainly not in Albania.

Helicigona skipetaricus SUBAI 1995

Helicigona subzonata (MOUSSON 1858)
Syn./ssp.: pindica

Helicigona zebiana (STURANY 1907)

Codringtonia neocrassa ZILCH 1952
 

Helicinae
Former generic names for Helicinae: Acavus, Archelix, Coenatoria, Cornu, Euparypha, Helicogena, Iberus, Macularia, Otala, Pomatia, Tachea, Xerophila.
Subfamily systematics:
Genus Helix: KNIPPER 1939

Theba pisana (O. F. MÜLLER 1774)

Eobania vermiculata (O. F. MÜLLER 1774)

Cepaea vindobonensis (FÉRUSSAC 1821)
Syn./ssp.: austriaca, balcanica, expallescens, nigra, pallescens

Helix aspersa O. F. MÜLLER 1774
Syn./ssp.: adspersa, Cryptomphalus aspersus

Helix cincta MÜLLER 1774
Note: Occurence in Albania needs confirmation.

Helix dormitoris (KOBELT 1898)
Syn./ssp.: arnautorum, kolaschinensis

Helix lucorum LINNÉ 1758
Syn./ssp.: onixiomicra, onixiomigra, presbensis KOBELT 1905

Helix oestreichi (KOBELT 1902)
Note: Systematic position not clear. Belongs probably to Helix secernenda or to Helix philibinensis ROSSMÄSSLER 1839.

Helix pomatia LINNÉ 1758
Note: does not live in today's Albania.

Helix secernenda ROSSMÄSSLER 1847
Syn./ssp.: albanica WOHLBEREDT 1907 [a], edlaueri, inflata, ligata, pomatia sensu HAX.1974
Note: The systematics of H. secernenda should be revised.

Helix vladika (KOBELT 1898)
Syn./ssp.: vladica, wohlberedti
 

Bivalvia: Unionida
Unionoidea
Unionidae
Former generic names: Mytilus.
Family systematics: according to HAAS 1969, FALKNER 1990.
Genus Anodonta: GLÖER, P. & MEIER-BROOK, C. (1994): Süsswassermollusken. Ein Bestimmungsschlüssel für die Bundesrepublik Deutschland. 11. Auflage. -- 136 pp. Hamburg

Anodonta anatina (LINNÉ 1758)
Syn./ssp.: piscinalis

Anodonta cygnea (LINNÉ 1758)
Syn./ssp.: cellensis, deplanata, subcircularis, zellensis GMELIN 1791
Note: The taxon cellensis SCHRÖTER 1779 does not exist.

Anodonta cymbalica DROUET 1879
Note: Actual systematic position not known.

Microcondylaea compressa (MENKE 1830)
Syn./ssp.: bonellii, depressa

Unio crassus PHILIPSSON 1788
Syn./ssp.: carneus, pruinosus

Unio elongatulus C. PFEIFFER 1825
Syn./ssp.: decipiens DROUET 1881, fiscallianus, fiscallinus, pallens

Unio mancus LAMARCK 1819
Syn./ssp.: elongatus C. PFEIFFER

Unio pictorum (LINNÉ 1758)

Unio tumidus PHILIPSSON 1788
Note: Lives in Lake Shkodra.
 

Veneroida
Sphaerioidea
Sphaeriidae
Former generic names: Sphaerium.

Musculium lacustre (O. F. MÜLLER 1774)
 Pisidium casertanum (POLI 1791)

Pisidium subtruncatum MALM 1855
Note: Lives in Lake Ohrid.

Sphaerium parenzani (GAMBETTA 1930)
Note: Systematic position not clear.
 

Dreissenoidea
Dreissenidae
Former generic names: Dreissensia .

Dreissena blanci WESTERLUND 1890

Dreissena polymorpha (PALLAS 1771)
 

List of Lake Ohrid species
 

The following species are mentioned in the literature for the Ohrid basin.

Gastropoda:
Theodoxus fluviatilis (LINNÉ 1758)
Viviparus viviparus (LINNÉ 1758)
Valvata stenotrema POLINSKI 1929
Valvata rhabdota STURANY 1894
Valvata hirsutecostata POLINSKI 1929
Orientalina curta (KÜSTER 1852)
Ohridohoratia pygmaea (WESTERLUND 1902)
Ohridohoratia carinata (RADOMAN 1956)
Polinskiola polinskii (RADOMAN 1960)
Polinskiola sturanyi (WESTERLUND 1902)
Ohridohauffenia depressa (RADOMAN 1956)
Ohridohauffenia sublitoralis (RADOMAN 1962)
Ohridohauffenia rotonda (RADOMAN 1964)
Ohridohauffenia drimica (RADOMAN 1964)
Ohridohauffenia minuta (RADOMAN 1955)
Ohridohauffenia sanctinaumi (RADOMAN 1964)
Ohrigocea samuili HADZISCE 1956
Ohrigocea karevi HADZISCE 1956
Ohrigocea miladinovorum HADZISCE 1956
Ohrigocea stankovici HADZISCE 1956
Dolapia ornata (RADOMAN 1956)
Gocea ohridana HADZISCE 1956
Pseudohoratia ohridana (POLINSKI 1929)
Pseudohoratia brusinae (RADOMAN 1953)
Pseudohoratia lacustris (RADOMAN 1964)
Lyhnidia gjorgjevici HADZISCE 1956
Lyhnidia hadzii HADZISCE 1956
Lyhnidia karamani HADZISCE 1956
Lyhnidia stankovici HADZISCE 1956
Lyhnidia sublitoralis (RADOMAN 1967)
Strugia ohridana RADOMAN 1973
Zaumia kusceri (HADZISCE 1956)
Zaumia sanctizaumi (RADOMAN 1964)
Pyrgohydrobia grochmalickii (POLINSKI 1929)
Pyrgohydrobia sanctinaumi RADOMAN 1955
Pyrgohydrobia jablanicensis RADOMAN 1955
Chilopyrgula sturanyi (BRUSINA 1896)
Neofossarulus stankovici POLINSKI 1929
Macedopyrgula pavlovici (POLINSKI 1929)
Macedopyrgula wagneri (POLINSKI 1929)
Stankovicia baicaliiformis POLINSKI 1929
Trachyohridia filocincta (POLINSKI 1929)
Ohridopyrgula macedonica (BRUSINA 1896)
Ginaia munda (STURANY 1894)
Xestopyrgula dybowskii (POLINSKI 1929)
Micropyrgula stankovici (POLINSKI 1929)
Holandriana holandrii (C. PFEIFFER 1828)
Acroloxus lacustris (LINNÉ 1758)
Acroloxus macedonicus HADZISCE 1959
Acroloxus improvisus POLINSKI 1929
Aplexa hypnorum (LINNÉ 1758)
Ancylus fluviatilis O. F. MÜLLER 1774
Ancylus lapicidus HUBENDICK 1960
Ancylus scalariniformis STANKOVIC & RADOMAN 1953
Ancylus tapirulus POLINSKI 1929
Bathyomphalus contortus (LINNÉ 1758)
Planorbarius corneus (LINNÉ 1758)
Planorbis planorbis (LINNÉ 1758)
Planorbis macedonicus STURANY 1894
Gyraulus lychnidicus HESSE 1928
Gyraulus trapezoides POLINSKI 1929
Gyraulus albidus RADOMAN 1953
Gyraulus crenophilus HUBENDICK & RADOMAN 1959
Gyraulus fontinalis HUBENDICK & RADOMAN 1959
Hippeutis complanatus (LINNÉ 1758)
Galba truncatula (O. F. MÜLLER 1774)
Lymnaea stagnalis (LINNÉ 1758)
Radix relicta POLINSKI 1929
Radix peregra (O. F. MÜLLER 1774)
Radix auricularia (LINNÉ 1758)
Stagnicola palustris (O. F. MÜLLER 1774)

Bivalvia:
Microcondylaea compressa (MENKE 1830)
Unio crassus PHILIPSSON 1788
Pisidium amnicum (O. F. MÜLLER 1774)
Pisidium casertanum (POLI 1791)
Pisidium edlaueri KUIPER 1960
Pisidium henslowanum (SHEPPARD 1823)
Pisidium moitessierianum PALADILHE 1866
Pisidium personatum MALM 1855
Pisidium subtruncatum MALM 1855
Pisidium tenuilineatum STELFOX 1918
Sphaerium parenzani (GAMBETTA 1930)
Dreissena polymorpha (PALLAS 1771)
 

List of Lake Shkodra species
 

The following molluscs live in Lake Shkodra. The list has been compiled by Dh. DHORA.

Gastropoda:
Theodoxus fluviatilis (LINNÉ 1758)
Viviparus mamillatus (KÜSTER 1852)
Valvata piscinalis (O. F. MÜLLER 1774)
Valvata cristata O. F. MÜLLER 1774
Anagastina vidrovani RADOMAN 1973
Anagastina scutarica RADOMAN 1973
Anagastina gluhodolica RADOMAN 1973
Anagastina matjasici BOLE 1961
Anagastina urbana RADOMAN 1973
Anagastina zetaevallis RADOMAN 1973
Pyrgula annulata (LINNÉ 1758)
Bithynia tentaculata (LINNÉ 1758)
Holandriana holandrii (C. PFEIFFER 1828)
Physella acuta (DRAPARNAUD 1805)
Gyraulus albus (O. F. MÜLLER 1774)
Planorbis planorbis (LINNÉ 1758)
Planorbis carinatus O. F. MÜLLER 1774
Planorbarius corneus (LINNÉ 1758)
Galba truncatula (O. F. MÜLLER 1774)
Lymnaea stagnalis (LINNÉ 1758)
Omphiscola glabra (O. F. MÜLLER 1774)
Radix ovata (DRAPARNAUD 1805)
Radix peregra (O. F. MÜLLER 1774)
Stagnicola palustris (O. F. MÜLLER 1774)
Stagnicola corvus (GMELIN 1791)
Radix auricularia (LINNÉ 1758)
Radix ampla (HARTMANN 1821)-complex

Bivalvia:
Unio crassus PHILIPSSON 1788
Unio elongatulus C. PFEIFFER 1825
Unio mancus LAMARCK 1819
Unio pictorum (LINNÉ 1758)
Unio tumidus PHILIPSSON 1788
Anodonta anatina (LINNÉ 1758)
Anodonta cygnea (LINNÉ 1758)
Microcondylaea compressa (MENKE 1830)
Musculium lacustre (O. F. MÜLLER 1774)
Dreissena blanci WESTERLUND 1890
 

Albanian landsnails. In the background the castle of Shkodër.     Photo by V. Wiese, Haus der Natur Cismar.
1 - Poiretia cornea, Pomatias elegans, Chondrula tridens, Zebrina detrita, Lindholmiola girva, Eobania vermiculata.
2 - Cochlostoma roseoli, Granaria frumentum, Strigilodelima conspersa, Montenegrina laxa, Delima montenegrina.
3 - Oxychilus glaber, Cernuella virgata, Monacha cartusiana, Xeromunda vulgarissima, Helicigona setigera.

This page is managed by Francisco Welter-Schultes (homepage). I am working in the Zoological Institute of Göttingen University. I receive a doctoral scholarship for Albinaria research from the Evangelisches Studienwerk.
I am always interested in new literature on Albanian molluscs.

Last modified 2.10.2002

      Montenegrina subcristata

Address of the first author of this list:
Dr. Dhimitër Dhora,  Universiteti i Shkodrës "Luigj Gurakuqi",  Departamenti i Biologjisë,  Shkodër, Albania

muell@schrottmails.de hier klicken